Born Middleton in Nova Scotia 1st May 1882. BSc at Mount Allison University 1899-1903. PhD, University of Chicago, 1908. Lecturer in Biology, St Thomas's Hospital, London, 1912-1914. Lecturer in Cytology, Bedford College, London, 1912 and on Heredity in Relation to Cytology, Oxford University, 1914. Instructor in aerial gunnery, Royal Flying Corps, 1917-1918. Reader in Botany, University of London King's College, 1919?-1921. Professor of Botany, University of London King's College, 1921-1942. Emeritus Professor, King's College London, 1943. Research Fellow in Biology, Harvard University, 1946-1950. Married Laura Greer, 1955. Died 12th August 1962.
From the description of The Ruggles Gates Collection. ca. 1900-1962. (King's College London). WorldCat record id: 681144194