Dale E. Madden received the Ph. D (1954) in Animal Science from Iowa State College (University). As a student at Iowa State, he attended several classes taught by Professor Jay L. Lush. Lush was a professor of Animal Husbandry at Iowa State (1930-1971). He developed improvement of livestock through his genetic and biometric research and also founded the field of population genetics, which redefined the concept of animal breeds along statistical norms.
From the description of Papers, 1952. (Iowa State University). WorldCat record id: 55113278